Dark Souls Science - Not All Ultragreatswords Are Created Equal
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In this video, we visit the topic of ultragreatsword swing speeds. The long and short of it is that the Zweihander and Greatsword are equally fast, but faster than the other three ultras (Dragon Greatsword, Black Knight Greatsword, and Demon Great Machete). By "faster", I mean they have a shorter recovery time (thanks to commenter LuckySharm for pointing this out).
Moreover, we find that the upgrade level of the weapon doesn't affect the recovery time or swing speed.
If you don't want to watch through all of the +0/+15 comparisons (which are included for completeness, since they don't show any differences), you can skip to 6:48.
The YouTube user auspiciously named Balder Knight has a comparison of greatsword speeds: https://www.youtube.com/watch?v=KMVzEIl_rs0.